1. Bioproduction Research Institute, National Institute of Advanced and Industrial Science and Technology, Tsukuba, Ibaraki, Japan
2. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
3. Microbiology Research Center for Sustainability, University of Tsukuba, Tsukuba, Ibaraki, Japan
4. Biotechnology Research Center, The University of Tokyo, Tokyo, Japan