1. INRA, UMR1253 STLO, F-35000 Rennes, France
2. Agrocampus Rennes, UMR1253, F-35000 Rennes, France
3. Centre for Infectious Diseases, The Chancellor's Building, New Royal Infirmary, University of Edinburgh, Edinburgh EH16 4SB, Scotland, United Kingdom
4. Research Technologies Branch, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Diseases, The National Institutes of Health, Hamilton, Montana 59840
5. Laboratory of Human Bacterial Pathogenesis, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Diseases, The National Institute of Health, Hamilton, Montana 59840