1. Institute of Hydrobiology, Chinese Academy of Sciences , Wuhan, China
2. University of Chinese Academy of Sciences , Beijing, China
3. College of Fisheries and Life Science, Dalian Ocean University , Dalian, China
4. State Key Laboratory of Freshwater Ecology and Biotechnology, The Key Laboratory of Breeding Biotechnology and Sustainable Aquaculture, Hubei Hongshan Laboratory, the Innovation Academy of Seed Design, Chinese Academy of Sciences , Wuhan, China