1. Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton CB10 1SA, United Kingdom
2. Department of Biological Sciences and Pest Management, Colorado State University, Fort Collins, Colorado 80523
3. Department of Ecology, Evolution and Behavior, University of Minnesota, St. Paul, Minnesota 55108
4. Department of Horticulture and Crop Science Ohio Agricultural Research and Development Center, The Ohio State University, Wooster, Ohio 44691
5. Department of Plant Pathology, University of Minnesota, St. Paul, Minnesota 55108