1. Cooperative Institute for Research in Environmental Sciences, University of Colorado, Boulder, Colorado, USA
2. Departamento de Biología y Geología, Física y Química Inorgánica, Escuela Superior de Ciencias Experimentales, Universidad Rey Juan Carlos, Móstoles, Spain
3. Institute of Arctic and Alpine Research, University of Colorado, Boulder, Colorado, USA
4. Environmental Studies Program, University of Colorado, Boulder, Colorado, USA
5. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA