1. Division of Cell and Developmental Biology, Wellcome Trust Biocentre, University of Dundee, Dundee DD1 5EH, United Kingdom
2. Cancer Research UK, Molecular Pharmacology Unit, Biomedical Research Center, Ninewells Hospital and Medical School, Dundee DD1 9SY, United Kingdom
3. Division of Biological Chemistry, Wellcome Trust Biocentre, University of Dundee, Dundee DD1 5EH, United Kingdom
4. Division of Applied Medicine, IMS Building, Foresterhill, Aberdeen AB25 2ZD, United Kingdom
5. Roslin Institute, Edinburgh EH25 9PS, United Kingdom
6. Institute of Basic Medical Sciences, University of Oslo, Oslo, Norway