1. Pearl River Fishery Research Institute, Chinese Academy of Fishery Sciences, Key Laboratory of Fishery Drug Development, Ministry of Agriculture, Key Laboratory of Aquatic Animal Immune Technology, Guangzhou, People's Republic of China
2. Department of Animal Sciences, University of Florida, Gainesville, Florida, USA
3. State Key Laboratory of Agricultural Microbiology, Huazhong Agricultural University, Wuhan, Hubei, People's Republic of China
4. College of Fisheries, Huazhong Agricultural University, Wuhan, Hubei, People's Republic of China
5. Department of Infectious Diseases and Pathology, University of Florida, Gainesville, Florida, USA