1. Department of Bioscience and Biotechnology, University of the Ryukyus, Senbaru, Nishihara, Okinawa, Japan
2. Graduate School of Science, Osaka Prefecture University, Gakuencho, Sakai, Osaka, Japan
3. Biomedical Research Institute, National Institute of Advanced Industrial Science and Technology, Higashi, Tsukuba, Ibaraki, Japan