1. Cell and Developmental Biology, Biological Sciences, University of California, San Diego, 9500 Gilman Drive, La Jolla California 92093-0368
2. Department of Developmental Biology and Neurosciences, Graduate School of Life Sciences, Tohoku University, Aoba, Sendai 980-8578, Japan
3. Biochemistry Department, Oxford University, South Parks Road, Oxford OX1 3QU, United Kingdom