1. Laboratory of Bioresource and Technology, Department of Integrated Biosciences, Graduate School of Frontier Sciences, The University of Tokyo, Kahiwa, Chiba, Japan
2. Laboratory of Plant Virology, Department of Applied Biological Sciences, Faculty of Agriculture, Saga University, Honjo-machi, Saga, Japan
3. The United Graduate School of Agricultural Sciences, Kagoshima University, Kagoshima, Japan