1. Laboratory for MetaSystems Research, RIKEN Quantitative Biology Center, Yokohama 230-0045, Japan
2. Present address: Department of Biology, Indian Institute of Science Education and Research (IISER), Bhopal, India.
3. Graduate School of Frontier Sciences, University of Tokyo, Kashiwa, Chiba 277-8561, Japan
4. Present address: RIKEN Research Center for Allergy and Immunology, Yokohama 230-0045, Japan.
5. Institute of Cell Biology, School of Biological Sciences, University of Edinburgh, Mayfield Road, Edinburgh EH9 3JR, United Kingdom