1. Department of Land Resource Science, University of Guelph, Guelph, Ontario N1G 2W1, Canada
2. Department of Earth Sciences, University of Ottawa, Ottawa, Ontario K1N 6N5, Canada
3. Argonne National Laboratory, Argonne, Illinois 60439
4. Department of Molecular and Cellular Biology, University of Guelph, Guelph, Ontario N1G 2W1, Canada