1. Institute for Animal Health, Ash Road, Pirbright, Surrey GU24 0NF, United Kingdom
2. Division of Environmental and Evolutionary Biology, University of Glasgow, Glasgow G12 8QQ, United Kingdom
3. Met Office, FitzRoy Road, Exeter, Devon EX1 3PB, United Kingdom
4. Animal Health and Welfare, Defra, 1a Page Street, London SW1P 4PQ, United Kingdom