1. Department of Virology, Nagoya University Graduate School of Medicine, Nagoya, Aichi, Japan
2. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), Kawaguchi, Saitama, Japan
3. Department of Virology and Parasitology, Fujita Health University School of Medicine, Toyoake, Aichi, Japan