1. Institute of Virology and AIDS Research, First Hospital of Jilin University, Changchun, Jilin, China
2. Key Laboratory of Zoonosis, Ministry of Education, Institute of Zoonosis, Jilin University, Changchun, Jilin, China
3. Jilin Provincial Center for Disease Control and Prevention, Changchun, Jilin, China
4. Department of Molecular Microbiology and Immunology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, USA