1. MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, John Radcliffe Hospital, Oxford OX3 9DS, United Kingdom
2. Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, United Kingdom
3. MRC Bioinformatics Capacity Development Research Unit, South African National Bioinformatics Institute, University of the Western Cape, Cape Town, South Africa
4. Institute of Evolutionary Biology, University of Edinburgh, Edinburgh EH9 3JT, United Kingdom
5. Department of Infectious Diseases, King's College London School of Medicine, London SE1 9RT, United Kingdom