1. Department of Bioresource Science, Ibaraki University College of Agriculture, Ami-machi, Ibaraki, Japan
2. Center for Conservation Science and Restoration Techniques, National Research Institute for Cultural Properties, Tokyo, Taito-ku, Tokyo, Japan
3. Department of Computational Biology, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwanoha, Kashiwa, Chiba, Japan
4. Graduate School of Life and Environmental Science, University of Tsukuba, Tennodai, Tsukuba, Ibaraki, Japan