1. Graduate School of Natural Science and Technology and Department of Biology, Faculty of Science, Okayama University, Tsushima-Naka 3-1-1, Kita-ku, Okayama 700-8530, Japan
2. Graduate School of Biosphere Science and Faculty of Applied Biological Science, Hiroshima University, Kagamiyama 1-4-4, Higashi-Hiroshima, Hiroshima 739-8528, Japan