1. Department of Pathology, Microbiology, & Immunology, Vanderbilt University School of Medicine, Nashville, Tennessee, USA
2. Department of Veterinary Medical Science, University of Miyazaki, Miyazaki, Japan
3. Center for Animal Disease Control, University of Miyazaki, Miyazaki, Japan
4. Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, USA
5. Veterans Affairs Tennessee Valley Healthcare System, Nashville, Tennessee, USA