1. School of Minerals Processing and Bioengineering, Central South University , Changsha, China
2. Key Laboratory of Biometallurgy of Ministry of Education, Central South University , Changsha, China
3. Hunan Tobacco Science Institute , Changsha, China
4. University of Chinese Academy of Sciences , Beijing, China
5. State Key Laboratory of Microbial Resources, Institute of Microbiology, Chinese Academy of Sciences , Beijing, China