1. Ocean EcoSystems Biology Unit, RD3, GEOMAR Helmholtz Centre for Ocean Research Kiel, Kiel, Germany
2. Kiel University, Kiel, Germany
3. Department of Microbiology, Oregon State University, Corvallis, Oregon, USA
4. Marine Biological Laboratory, Woods Hole, Massachusetts, USA
5. Max Planck Institute for Evolutionary Biology, Plön, Germany