1. Department of Cell and Developmental Biology, Vanderbilt University, Nashville, Tennessee, USA
2. Department of Pediatrics, Children’s Hospital of Pittsburgh, University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, USA
3. Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, USA
4. Epithelial Biology Center, Vanderbilt University, Nashville, Tennessee, USA
5. Department of Molecular Physiology and Biophysics, Vanderbilt University, Nashville, Tennessee, USA
6. Department of Pathology, Microbiology, and Immunology, Vanderbilt University School of Medicine, Nashville, Tennessee, USA
7. Veterans Affairs Tennessee Valley Healthcare System, Nashville, Tennessee, USA
8. Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, USA
9. Department of Cell and Developmental Biology, University of Michigan, Ann Arbor, Michigan, USA