1. School of Earth, Atmospheric and Environmental Sciences; Williamson Building, University of Manchester; Manchester M13 9PL United Kingdom
2. Office of Biological and Environmental Research SC-75 / Germantown Building; U.S. Department of Energy; 1000 Independence Ave., S.W. Washington DC 20585-1290
3. School of Biosciences; The University of Birmingham; Birmingham B15 2TT United Kingdom