1. Department of Biological Sciences; University of Essex; Colchester C04 3SQ United Kingdom
2. Enterovirus Laboratory; KTL, Mannerheimintie 166 00300 Helsinki Finland
3. Institute for Animal Health (IAH); Pirbright Laboratory; Ash Road, Pirbright, Woking Surrey GU24 ONE United Kingdom
4. Department of Virology; Haartman Institute, University of Helsinki; P.O. Box 21 00014 Helsinki Finland