1. Graduate School of Life Sciences, Toyo University, Itakura-machi, Gunma, Japan
2. Bio Resilience Research Center, Toyo University, Itakura-machi, Gunma, Japan
3. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
4. Microbiology Research Center for Sustainability (MiCS), University of Tsukuba, Tsukuba, Ibaraki, Japan