1. Department of Biology, Haverford College, Haverford, Pennsylvania, USA
2. Department of Genome Sciences, University of Washington, Seattle, Washington, USA
3. College of Earth, Ocean, and Atmospheric Sciences, Oregon State University, Corvallis, Oregon, USA
4. Marine Chemistry and Geochemistry, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, USA
5. Department of Biological Sciences, University of New Hampshire, Durham, New Hampshire, USA