1. Department of Pharmaceutical Sciences, School of Pharmacy, University of Maryland , Baltimore, Maryland, USA
2. Department of Microbiology, Immunology, and Cell Biology, West Virginia University , Morgantown, West Virginia, USA
3. Vaccine Development Center at West Virginia University Health Sciences Center , Morgantown, West Virginia, USA
4. Department of Microbiology and Immunology, School of Medicine, University of Maryland , Baltimore, Maryland, USA