1. Division of Virology, Institute of Medical Science, University of Tokyo, Tokyo, Japan
2. Center for Global Viral Diseases, National Center for Global Health and Medicine, Tokyo, Japan
3. Shinwa Corporation, Tokyo, Japan
4. Department of Special Pathogens, International Research Center for Infectious Diseases, Institute of Medical Science, University of Tokyo, Tokyo, Japan
5. Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin—Madison, Madison, Wisconsin, USA