1. CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China
2. Marine Environmental Engineering Center, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China
3. University of Chinese Academy of Sciences, Beijing, China
4. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China