1. Department of Biotechnology and Environmental Biology, RMIT University, Bundoora West Campus, Bundoora, Melbourne, Victoria 3083, Australia
2. University of Melbourne, Rural Veterinary Unit, Maffra, Victoria 3860, Australia
3. Institute for Animal Health, Compton Laboratory, Compton RG20 7NN, United Kingdom
4. School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, Leicestershire LE12 5RD, United Kingdom