1. Department of Earth, Atmospheric, and Planetary Sciences, Massachusetts Institute of Technology, Cambridge, Massachusetts, USA
2. Department of Marine Chemistry and Geochemistry, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, USA
3. Microbial and Environmental Genomics, J. Craig Venter Institute, San Diego, California, USA
4. Marine Biology Research Division, Scripps Institution of Oceanography, La Jolla, California, USA