1. Université Paris-Saclay, CNRS, AgroParisTech, Ecologie Systématique et Evolution, Orsay, France
2. Université Paris-Saclay, CEA, CNRS, Institute for Integrative Biology of the Cell (I2BC), Gif-sur-Yvette, France
3. Laboratory of Biodiversity and Valorization of Arid Areas Bioresources (BVBAA), Faculty of Sciences, Gabès, Tunisia