1. Division of Molecular Epidemiology, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan
2. Sukra Raj Tropical and Infectious Disease Hospital, Kathmandu, Nepal
3. Department of Microbiology, Infectious and Tropical Disease Research and Prevention Center, Kathmandu, Nepal
4. Department of Medical Microbiology and Genitourinary Medicine, University of Liverpool, Liverpool, United Kingdom
5. Liverpool School of Tropical Medicine, Liverpool, United Kingdom