1. Institute of Applied Biochemistry, University of Tsukuba, Tsukuba, Ibaraki 305-8572
2. Department of Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo-ku, Tokyo 113-8657
3. Department of Applied Biological Sciences, College of Bioresource Sciences, Nihon University, Fujisawa, Kanagawa 252-8510, Japan