1. Department of Molecular Genetics, University of Toronto, 1 King's College Circle, Toronto, Ontario M5S 1A8, Canada
2. Program in Developmental and Stem Cell Biology, Research Institute, The Hospital for Sick Children, 101 College Street, Toronto, Ontario M5G 1L7, Canada
3. Department of Biochemistry, University of Toronto, 1 King's College Circle, Toronto, Ontario M5S 1A8, Canada