1. Department of Bioscience, Tokyo University of Agriculture, Tokyo, Japan
2. Genome Research Center, NODAI Research Institute, Tokyo University of Agriculture, Tokyo, Japan
3. Graduate School of Frontier Sciences, The University of Tokyo, Chiba, Japan
4. Laboratory of Microbial Technology, Division of Applied Molecular Microbiology and Biomass Chemistry, Department of Bioscience and Biotechnology, Faculty of Agriculture, Graduate School, Kyushu University, Fukuoka, Japan
5. Department of Environmental Science, Musashino University, Tokyo, Japan