1. Department of Water Resources Management, Faculty of Marine and Fisheries Sciences, Raja Ali Haji Maritime University, Jl. Senggarang Polytechnic. Tanjungpinang, Riau Islands 29111, Indonesia
2. Laboratorium Marine Biotechnology, Fakultas Ilmu Kelautan dan Perikanan, Universitas Maritim Raja Ali Haji, Tanjungpinang, Indonesia