1. Okinawa Churashima Research Center, 888 Ishikawa, Motobu, Okinawa 905–0206, JAPAN
2. Graduate School of Engineering and Science, University of the Ryukyus, Nishihara, Okinawa 903–0213, JAPAN
3. Tropical Biosphere Research Center, University of the Ryukyus, Nishihara, Okinawa 903–0213, JAPAN