1. Faculty of Agriculture, Forestry, and Food Engineering, Yibin University, Yibin 644007, China
2. State Key Laboratory of Genetic Resources and Evolution, and Yunnan Key Laboratory of Biodiversity and Ecological Security of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China
3. Central South Inventory and Planning Institute of National Forestry and Grassland Administration, Changsha 410014, China