1. Department of Biology, Graduate School of Science, Toho University, Miyama 2–2–1, Funabashi, Chiba 274–8510, JAPAN
2. Chiba Wildlife Research Society, Semata 962–40, Ichihara, Chiba 290–0151, JAPAN
3. Department of Biology, Faculty of Science, Toho University, Miyama 2–2–1, Funabashi, Chiba 274–8510, JAPAN