1. Research and Education Center for Natural Sciences, Keio University, Hiyoshi, Kohoku-ku, Yokohama, Kanagawa 223–8521, JAPAN
2. Department of Biology, Keio University, Hiyoshi, Kohoku-ku, Yokohama, Kanagawa 223–8521, JAPAN
3. School of Biology, Tokai University, Minamisawa Minami-ku, Sapporo, Hokkaido 005–8601, JAPAN
4. Faculty of Agriculture, Meiji University, Higashimita, Tama-ku, Kawasaki, Kanagawa 214–8571, JAPAN