1. Key Laboratory of Marine Environment and Ecology, Ministry of Education, Ocean University of China, Qingdao 266100, P. R. China yonggang@ouc.edu.cn
2. College of Environmental Science and Engineering, Ocean University of China, Qingdao 266100, P. R. China lihonglei200401@yahoo.cn
3. First Institute of Oceanography, State Oceanic Administration, Qingdao 266061, P. R. China