1. Fisheries and Oceans Canada, Pacific Science Enterprise Centre, West Vancouver, BC V7V 1N6, Canada
2. Fisheries and Oceans Canada, Institute of Ocean Sciences, Sidney, BC V8L 5T5, Canada
3. Lorax Environmental Services, Vancouver, BC V6J 3H9, Canada
4. National Oceanography Centre, University of Southampton, Southampton SO14 3ZH, England, U.K.