1. Department of Agrobiological Science, Faculty of Agriculture, Ehime University, Matsuyama 790-8566, Ehime, Japan
2. Department of Microbiology, Faculty of Medicine, Oita University, Yufu, Oita 879-5593, Japan
3. School of Neuroscience, Virginia Polytechnic Institute and State University, 24061, Blacksburg, Virginia, United States