1. Department of Applied Life Sciences, Faculty of Agriculture, Shizuoka University, 836 Ohya, Shizuoka City, Shizuka 422-8529, Japan
2. Department of Biological Sciences, Faculty of Science, Hokkaido University, Kita 10 Nishi 8, Kita-ku, Sapporo, Hokkaido 060-0810, Japan
3. Oki Marine Biological Station, Education and Research Center for Biological Resources, Faculty of Life and Environmental Science, Shimane University, Oki, Shimane 685-0024, Japan