1. Laboratory of Animal Breeding and Genetics, Graduate School of Biosphere Science, Hiroshima University, Japan
2. Laboratory of Animal Genetics, Graduate School of Bioagricultural Sciences, Nagoya University, Japan
3. Japanese Avian Bioresource Project Research Center, Hiroshima University, Japan
4. Present address: College of Agriculture, Ibaraki University, Japan
5. Present address: Institute de Sélection Animale B.V., The Netherlands-EU