1. 1Maxwell H. Gluck Equine Research Center, Department of Veterinary Science, University of K entucky, Lexington, KUSA
2. 2INRS Centre Armand-Frappier et Santé Biotechnologique, Laval, Québec, Canada
3. 3Université Paris-Saclay, UVSQ, INRAE, BREED, Jouy-en-Josas, France
4. 4Ecole Nationale Vétérinaire d’Alfort, BREED, Maisons-Alfort, France
5. 5Comparative Biomedical Sciences, Royal Veterinary College, Hatfield, UK