1. Computational Neurobiology Laboratory, The Salk Institute for Biological Studies, 10010 North Torrey Pines Road, La Jolla, California 92037, USA
2. Neurosciences Graduate Program, University of California San Diego, La Jolla, California 92093, USA
3. Institute for Neural Computation, University of California San Diego, La Jolla, California 92093, USA
4. Division of Biological Sciences, University of California San Diego, La Jolla, California 92093, USA
5. CORIA, Rouen Normandie Université, Campus Universitaire du Madrillet, F-76800 Saint-Etienne du Rouvray, France