1. Institute of Industrial Science, The University of Tokyo, 4-6-1 Komaba, Meguro-ku, Tokyo 153-8505, Japan
2. Department of Mathematical Informatics, The University of Tokyo, Bunkyo-ku, Tokyo 113-8656, Japan
3. CREST, JST, 4-1-8 Honcho, Kawaguchi, Saitama 332-0012, Japan
4. Mathematical Neuroinformatics Group, Advanced Industrial Science and Technology, Tsukuba, Ibaraki 305-8568, Japan
5. Center for Research in Agricultural Genomics (CRAG), Consorci CSIC-IRTA-UAB-UB, Barcelona 08193, Spain