1. Institute for Integrated Cell-Material Sciences (WPI iCeMS), Kyoto University, 606-8501 Kyoto, Japan
2. Physical Chemistry of Biosystems, Institute of Physical Chemistry, University of Heidelberg, D69120 Heidelberg, Germany
3. Division of Systems Science, Department of Systems Innovation, Graduate School of Engineering Science, Osaka University, 560-8531 Toyonaka, Japan
4. Department of Physics, Graduate School of Science, Kyoto University, 606-8501 Kyoto, Japan
5. Molecular Evolution and Genomics, Centre for Organismal Studies (COS), University of Heidelberg, D69120 Heidelberg, Germany